1Department of Biology, Faculty of Science, Khon Kaen University, Khon Kaen 40002, Thailand.
2Faculty of Environment and Resource Studies, Mahasarakham University, Mahasarakham 44000, Thailand.
3Department of Botany and Pharmacognosy, Faculty of Pharmaceutical Sciences, Khon Kaen University, Khon Kaen 40002, Thailand.
4Department of Botany, University of Hawaii at Manoa, Hawaii 96822, USA.
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