1Hainan Key Laboratory of Tropical Oil Crops Biology , Coconut Research Institute, Chinese Academy
of Tropical Agricultural Sciences, Wenchang 571339, Hainan,China.
2The CAS/Shandong Provincial Key Laboratory of Coastal Environmental Process, Yantai Institute
of Coastal Zone Research, Chinese Academy of Sciences(CAS), Yantai 264003,Shandong, China.
3Institute for Life Sciences, Qingdao University of Science and Technology(QUST), Qingdao266042,
Email: [email protected], [email protected]