1School of Life Sciences, Lanzhou University, Lanzhou, 730000, PR, China.
2National Research Center for Conservation of Ancient Wall Paintings and Earthen Architecture Sites, Dunhuang, Gansu, 736200, PR, China.
3Key Scientific Research Base of Conservation for Ancient Mural, State Administration for Cultural Heritage, Dunhuang, Gansu 736200, PR, China.
4Conservation Institute of Dunhuang Academy, Dunhuang, Gansu, 736200, PR, China.
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