1Key Laboratory of Southwest China Wildlife Resources Conservation (Ministry of Education), College of Life Science, China West Normal University. No. 44 Yuying Road, Nanchong 637002, China.
2Key Laboratory for Biological Resource and Ecological Environment of the Ministry of Education, College of Life Sciences, Sichuan University. No. 29, Wangjiang Road, Chengdu 610064, P. R. China.
3College of Sciences, Tibet University, Lhasa 850000, China.
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